Nature Notes

by

Philip Radford

I expect that most of us are impressed by the large size of the eyes of dragonflies which, for the bigger species, seem to make up much of the head, with such a different appearance from that of a butterfly. Butterflies have long, clubbed antennae and relatively small eyes, while the antennae of dragonflies are short and often inconspicuous. In contrast, the antennae of moths are big and
often broad and feathery, with those of the males being much larger than those of the females; because of these sensitive antennae, some male moths are able to detect the presence of females , by scent, over sometim es quite remarkable distances. This long-range sensitivity is invaluable for night-flying moths where the population is low, at least if they are to have a chance of mating.
With certain moth species, the males fly by day while females only fly by night; an example here is the handsome Emperor Moth, with big eye spots on the wings, and which is found on the Quantock moors and on Exmoor. The small male Emperors, with big bushy antennae, fly on sunny days and are attracted, by scent, to females as they rest amongst heather. The thin, clubbed antennae of butterflies help the males to contact females by scent as well, although sight plays a part with these day-flying insects. Additionally, butterflies detect suitable flower scents using their antennae, so that the insect can have a feed of nectar; a butterfly feeds by inserting its proboscis, really a long sucking tube, into the chosen flower. Another use of antennae in butterflies is that, by scent, the correct plant can be detected on which the female can lay her eggs; often foot movements help release scent from a leaf.
If eggs are laid on the wrong plant, then hatched caterpillars are unable to feed.
Moths, like butterflies, feed on flower nectar but, for some species, this is only by night. However, one day-flying moth is the Humming-bird Hawk-moth and, as a migrant from Mediterranean lands, it can be seen, remarkably, flying in the sun in Quantock gardens; it is instructive to watch the hovering insect as it repeatedly inserts its long proboscis into suitable flowers. Last July I saw one of these moths which had evidently found a rich feeding source on a single red valerian, and was probing its proboscis into one multiple flower after another, and all on the same plant. The valerian was clearly ripe with tempting nectar; no doubt it was the sense of smell which guided the moth; there were other valerian plants around but they were all ignored Vision must be very different for butterflies, moths or dragonflies than for ourselves, although all the groups have colour vision of sorts. It seems that these insects, broadly, can detect ultra-violet light and so recognise colour patterns on other insects, and flowers at times, which would not be recognised by people.
The three insect classes all have compound eyes, ranging from about five thousand lens facets in good-sized butterflies to around twenty-five thousand in large dragonflies. These compound eyes enable the insects to detect movements around them, and to get a rough focus on nearby small objects. Even so, this form of vision allows the insect to see an approaching enemy and to react, either by flying off if perched or to take evasive action in the air.
Anyway, returning to dragonflies with their very large compound eyes, these insects feed only on other insects, which they catch in the air; flowers cannot be of any interest to dragonflies, except as perches. Sometimes dragonflies feed on other dragonflies, although not necessarily of the same species, and butterflies are often victims for the larger types; obviously, much aerial agility is
required for successful hunting by dragonflies, and so much depends on vision. Compound eyes essentially, have multiple facets, each having a lens on the surface and a cone-shaped lens beneath: ligh t rays are directed to a sensitive membrane, and the image is then relayed to the brain. The detailed structure of such an eye is cer tai nly remark able. With the large dragonflies , the eyes represent much of the head in size; in consequence, the insect can sense movement around it in front, at the side and even behind. In addition to these compound eyes, dragonflies have three light-sensitive spots (ocelli), each with one lens, on the top of the head; these simple eyes help the insect to keep stable in the air. The dragonflies' big eyes and small antennae must indicate how these insects live. Dragonflies catch their food and avoid enemies through vision; the short antennae give some sense of smell and help the insect to feel, which must be useful when eating its freshly-caught prey.